phrynoidis aspera (river toad, rough toad, giant asian toad, kodok buduk sungai, kodok puru besar)

amphibian animal asian asper aspera base besar black blunt body bony broad brown buduk bufo color connected covered crests dark derives females finger fish frog fully giant gland gray green head kodok large leaf length lizard longest males mm name nature nuptial ovoid pads parotoid phrynoidis puru reptile ridge river rough skin snake snout-vent species stout sungai sungal supraorbital supratympanic texture toad toe tortoise tubercles turtle tympanum warts webbed wildlife yellow