orange and brown butterfly wings open on a leaf

against and androconia antenna arthropod backdrop basking beasts blade brown butterfly chippenham closed clusters dimorphism elderberries elderberry euarthropoda eye eyespot fauna female field gate gatekeeper grass hedge her imago insect keeper leaf leaves lepidoptera maniola meadow mini nymphalidae open pyronia resting ripening satyrids satyrinae satyrines sexual spot sun tithonius tithonus unripe wildlife wiltshire wings