natterjack toad

ainsdale amphibia amphibian amplexus aquatic birkdale breeding bufo burrowing calamita coast coastal conservation conserve countryside coupling damp declining diurnal dune ecological ecology england female flooded habitat male mating natterjack natterjacks natural nature nightingale nocturnal north omnivorous pair pool pools population protected running seasonal sefton shallow slacks southport spawning spring stripe system toad toads two ugly uk warty west wetland wildlife yellow