male american toad (anaxyrus americanus) in wetland

male american toad anaxyrus americanus stands erect shallow wetland vocal sac partially inflated waits female mate toad stand posture erect tall forelimbs strong throat dark darkness outdoors creature wildlife macro close-up closeup wet wetland pond shallow wait waiting frog amphibian mating night bufonidae bufo anaxyrus americanus brown warts bumps bumpy nocturnal male female reproduction north america ohio environment environmental behavior ethology nature water animal lake