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howler monkeys high in a tree in gandoca manzanillo refuge, costa rica
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alouatta
alouattinae
are
costa
gandoca
genus
grandoca
high
holder
holwer
howler
largest
manzanillo
monkey
monkeys
monkies
monks
monotypic
native
new
refuge
rica
subfamily
tree
trees
tropical
tropics
wild
wildlife
world