howler monkey (alouatta palliata) in cano negro costa rica

howler monkey alouatta palliata resting tree branch caaplusmn negro wildlife refuge costa rica mammal primate nature tropical rainforest jungle forest habitat conservation biodiversity animal fauna flora sanctuary environment natural wild central america canopy arboreal foliage greenery landscape scenery tourism travel adventure exploration wildlifephotography ecotourism preservation protection species wilderness caño Cañonegro