emperor dragonfly (anax imperator) female's empty exuvia

dragonfly female emperor empty exuvia anax imperator dragonflies damsel fly damselflies skin insect animal nature species odonata moulting exoskeleton aquatic nymph metamorphosis life cycle closeup macro wildlife invertebrate biology entomology larval pond water reeds natural habitat environment emergence transformation larva shedding predator ecosystem exuviae history development leaf close insects