coprinus sp. - group of saprotrophic dung beetles on old horse manure, ukraine

coprinus agaricaceae agaricomycetes basidiomycota fungi mushroom saprotrophic dung manure horse old ukraine nature wildlife insects macro closeup ecology decomposition coprophagous scarab scarabaeidae entomology environment biodiversity ecosystem soil nutrient recycling detritivore agriculture field natural habitat behavior lifecycle species scavenger symbiosis mutualism coprophagy