coprinus sp. - group of saprotrophic dung beetles on old horse manure, ukraine

agaricaceae agaricomycetes agriculture basidiomycota behavior biodiversity closeup coprinus coprophagous coprophagy decomposition detritivore dung ecology ecosystem entomology environment field fungi habitat horse insects lifecycle macro manure mushroom mutualism natural nature nutrient old recycling saprotrophic scarab scarabaeidae scavenger soil species symbiosis ukraine wildlife