caloplaca verruculifera and other types of crustose lichens on a concrete wall of a pier near the black sea, odessa

lichen caloplaca verruculifera polycauliona verrucoplaca fungus teloschistaceae crustose saxicolous thallus apothecia odessa ukraine pier wall black coastal rock biodiversity ecology nature organism symbiosis mycology phycology teloschistales ascomycota mycobiota photobiont symbiont microhabitat epilithic bioindicator conservation species taxonomy morphology reproduction spore habitat distribution substrate colonization adaptation resilience weathering erosion biomonitoring